1/2] V; inner metatarsal tubercle distinct, small, elliptical; outer metatarsal tubercle absent; subarticular tubercles round, none bifid; supernumerary tubercles scattered on and around proximal ends of toes; terminal disc of toes elliptical, slightly wider than toes; terminal disc of Toes III, IV, and V slightly larger than those of Toes I and II; tarsal fold well-defined and flap-like.
1/2]V (UMMZ 239649); ulnar tubercles distinct, forming row; tarsal fold distinct; flanks and dorsum with same coloration and pigmentation pattern in preservative (Straughan & Wright 1969).
3) in females; inner tarsal fold
absent, inner tarsal surface bearing one tubercle, poorly defined; outer edge of tarsus without tubercles; knee and heel without prominent tubercles; inner metatarsal tubercle elongated, near 2-3 times size of rounded outer; supernumerary plantar tubercles numerous, poorly defined; subarticular tubercles large and prominent, rounded to oval; toes lacking narrow lateral fringes; no webbing; discs on all toes, discs of toes II-V expanded, larger than those of fingers; Toe V much longer that Toe III; tip of Toe V reaches to the distal border of the distal subarticular tubercle of Toe IV (condition C; Lynch and Duellman 1997).
mykter and based on our observations, discuss what are the conditions of the characters rostral keel, skin thickness, thoracic fold, and tarsal fold in this species.
3% of eye diameter), concealed dorsally by supratympanic fold and posteriorly by skin; tympanic ring distinct, slightly elevated anteriorly, not elevated posteriorly; thoracic fold absent; four or five irregular, shallow thoracic grooves present; thoracic grooves interrupted medially; arms covered with tubercles dorsally and ventrally; ventral tubercles forming low ridge along external edge of ulna; tarsal fold not evident; outer metatarsal tubercle present; webbing formula on foot I1-[2.
6 times the interorbital distance; (7) first finger equal or slightly longer than second; (8) disc on third finger more than 3/5 size of tympanum; (9) keels on fingers absent; (10) third finger not swollen in male; (11) short thick cloacal fold, with entire free border; (12) tarsal fold distinct, raised, not ending in tubercle; (13) inner metatarsal tubercle oval, about 2.
meridensis given within parentheses), by being a smaller frog (larger; see Table 1), and having the tip of snout more acute (rounded), tympanum with paler coloration (darker), larger dark brown dorsal spots (smaller), two irregular bands from upper eyelids to the level of shoulders (absent), oblique inguinal band larger (shorter), throat and chest with inconspicuous spots made up of fine dark stippling that do not form well-defined spots (having discrete dark brown spots), ventral surfaces of arms and thighs almost immaculate (well-differentiated dark brown little spots), tarsal fold evident (almost absent), metacarpal tubercle more pronounced (less pronounced), tend to have a more extended foot web (less webbed), and bearing conspicuous folding flaps on toes (bearing keels).
Diagnosis: (1) skin of dorsum shagreen with flat warts, that of venter areolate; short dorsolateral folds present; (2) tympanum round, its length 45-54 % that of eye; (3) snout short, subacuminate in dorsal view, rounded in profile; canthus rostralis sharp; (4) upper eyelid narrower than interorbital distance, no cranial crests; (5) vomerine odontophores oval; (6) vocal sac and nuptial pad present in males; (7) first finger shorter than second; fingers lack disks but with ventral pads; (8) fingers bear lateral keels; (9) ulnar tubercles present, fusing to form ridge; (10) round tubercle on heel; outer tarsal fold
present; short inner tarsal fold
; (11) inner metatarsal tubercle oval, 1.
6 times wider than adjacent phalanx; (7) fingers with thick lateral fringes; thickened edge along outer edge of fourth finger, from base of pad to close to outer metacarpal tubercle; (8) cloacal fold short; (9) tarsal fold
strong, not ending in tubercle but having a little distal enlargement; (10) foot-web formula: I2.
Conical tubercle on heel; outer edge of tarsus with row of subconical tubercles; short inner tarsal fold
on distal 1/4 of tarsus; inner metatarsal tubercle 2 1/2 times as long as wide, outer round, subconical, 1/4 size of inner; plantar surface bearing numerous supernumerary tubercles; basal subarticular tubercles longer than wide, distal ones round, all non-pungent; toes bearing lateral fringes but no webbing; all toes with ventral pads; disk of I not expanded, of II-V expanded (1 1/4-1 1/2 width of digit below disk); tip of toe III to distal edge of penultimate subarticular tubercle of toe IV, tip of toe V to 1/2 way between distal and penultimate subarticular tubercles of toe IV; heels overlapping when flexed hindlimbs held perpendicular to sagittal plane; shank 45.
lynchi and that species has a curved inner tarsal fold
Ford & Savage (1984:5) characterized the three frogs as having "narrow nonemarginate finger and toe disks, no tarsal fold
or tubercle, no toe webbing, finger I shorter than finger II, a strongly granulate (areolate) venter, a distinct sub-integumentary inguinal gland, and no vocal slits in adult males.