Tree 10-15 m tall, small young branches yellowish gray to purplish brown, mature branches yellowish gray to yellowish brown, Stipule caduceus, linear lanceolate, apex acuminate, young petiole densely hairy, soon hairless; leaf blade ovate to broadly ovate, glabrous or thickly hairy when young, soon glabrescent, base rounded or sub-cordate or sometime cordate, lustrous green above, margin long spinulose
serrate, apex shortly acuminate or long tapering acute apex.
Achenes 1.3-2.5 mm long, ovoid to globose, with obtuse, apiculate apex, irregularly transversely tuberculate and papillate, sometimes the papillae near the base elongate and spinulose
; hypogynium narrow, somewhat trigonous, with 6 finely puberulent globose tubercles in distinct pairs.
0.4 cm long, margins subdensely to sparsely spinulose
, spines 4-7 x 3-5 mm, 5-15 mm apart, castaneous-red near the apex.
4) and in other cottoids by Jackson (2004); spine, a sharp needle-shaped bony structure on a dermal tubercle or other ossicle (Taurulus); spinule, a tiny spine projecting from a larger spine or horn; spinulose
tubercle, a tubercle covered with many tiny spinules (Triglopsis); spiny rod, a spine-bearing fin ray of the paired fins (e.g.
Astrocaryum chambira differs from the other species of subgenus Astrocaryum in having the biggest fruit, with epicarp glabrate when unripe, usually smooth, sometimes sparsely spinulose
, greenish to yellowish when ripe, and a distinctive massive obconical perianth, and in having yellowish spines on the petioles of juvenile plants.
Microspores light gray to brown, papillose to spinulose
Carapace low, covered with yellowish brown thin hairs and black spinulose
Flowers with perianth of 1-5 slender bristles (sometimes 1 or 2 reduced or rudimentary), shorter than the achene or rarely overtopping the summit, retrorsely spinulose
nearly to the base, colorless to pinkish or reddish-brown; stamens 3; anthers (1.4-)2-3.2 mm long, yellow to orangebrown; styles bifid (rarely some trifid in some spikelets).
cantabrica Echinate hairs Present Absent Absent Staminal Glabrous Glabrous Hairy at base filaments Bract and Variable; at Basal Half way of the bracteoles base, middle or pedicel, rarely top of the basal pedicel Stigma Broadly Obconic to Obconic to capitate, capitate capitate disk-like Corolla length 2-3 3-3.5 3.5-5 (mm) Corolla lobes 0.4-0.8 (1.5) 1-1.5 0.8-1 (mm) Anther size 0.7-1.1 0.8-1 0.6-0.9 (mm) Anther 0.3-0.7 Toothed 0.2-0.4 0.5-0.7 appendages (mm) Spinulose
* Dryopteridaccae Dryopteris cartbusiana Spinulose
Wood Fern (Vill.) H.P.