Tergites sculptured with transverse striae except the area between median setae
on tergies II (Fig.
--Parafacial with a row of setulae on basal two-thirds, flanked by shorter setulae; postpronotal lobe with three postpronotal setae
almost in line and one weaker anterior postpronotal seta positioning between the middle and inner postpronotal setae
; surstylus in lateral view sharper (see Toma and Guimaraes 2000: Figs.
Male sixth through eighth abdominal segments completely retractable within the anterior part of the abdomen, gonocoxite and gonostylus of male terminalia extremely prolonged and thin; ovipositor elongated and protractible, cercus with simple setae
. Pupa: antennal horn with upper and lower spines, upper and lower frontal horn absent, upper cephalic margin thickened laterally; prothoracic spiracle and cephalic papillae long; abdominal dorsal segments with rows of small spicules, without spines.
Slide preparations were made as thin as possible to press setae
Coxa and basis fused, with 3 distal setae
. Endopod represented by 1 seta.
Williams and Hodges (1997) reported the presence of long femoral setae
in other species of Coccidae, i.e., Hemilecanium petasus (Hodgson, 1991) (as Etiennea petasus Hodgson, 1991), Kilifia americana Ben-Dov, 1979, Milviscutulus mangiferae (Green, 1889), Protopulvinaria pyriformis Cockerell, 1894 and Pseudokermes nitens (Cockerell, 1895), but did not specified on which of the three pairs of legs these very long setae
Orbital plate dark brown, the medial vertical setae
is closer to lateral vertical setae
and slightly toward the outer edge of the orbital plate, distance of or3 to or1, 82% (99%) of or3 to vtm (orbito-index, McEvey 1990), or1/or3 ratio 0.9 (paratype with or3 broken), or2/or1 ratio 0.5 (0.6), postocellar setae
62% (37%) and ocellar setae
88% (88%) of frontal length; vt index = 1.1 (0.9).
After examining specimens of the genera Nhandu Lucas, 1983, Proshapalopus Mello-Leitao, 1923, Pterinopelma and Vitalius Lucas, Silva & Bertani, 1993, a conspicuous tuft of setae
on the trochanters and coxae drew our attention.
Pterothorax: Trapezoidal, lateral margins slightly divergent, posterior margin convex, forming fine lateral ends; pteronotum not divided; pteronotal postero-lateral marginal setae
five to six on each side, arrangement 4,2+2,4 (4 latero-posterior and 2 posterior setae
); trichoid setae
present, thorn like setae
present; mesothoracic spiracles ventral in position; proepimeron expanded posteriorly and separated in median; mesofurcal pit present; meso-metasternal plate elongated and narrow, with one pair of normal fine seta on each.
Anepimeron with setae
. Tarsal claws toothed and similar in all legs, bent near base; empodium short, reaching bent in claws (Figure 8).
The most important character for the identification of male is spur prominent on the leg IV and bearing the pectinate setae
on the femur and trochanter.
The basal plate of the gills called the setobranch, was curved anteriorly and devoid of any gill filaments but have number of the basal spines and hair-like setobranch setae
towards their outer edge, while the apex and body (middle) parts have high number of filaments which were being arranged in rows across the gill axis and were originated from the anterior and posterior surface of the gill axis in body and apex, in addition to apical border of apex of the gill axis, while the lateral and medial surfaces of body and apex were devoid from any filaments.