The seed coats are moderately or deeply reticulate with layers of scalariform thickenings on the radial walls.
integra have nine layers of scalariform thickenings in the radial walls as compared to three in C.
Each cell has eight to 10 layers of scalariform thickenings, this character gives the seed a deep honeycomb appearance.
The combination of the size and shape of cells that make up the testa, the ornamentation of the radial walls of those cells, and the number of scalariform thickenings in the testa allow for precise identification at the species level.