There are only two genes of hydroxy cinnamoyl-CoA
quinate hydroxycinnamoyl transferase (HQT) and allene oxide synthase LjAOS were reported to correlated with the biosynthesis of chlorogenic acid during the floral organ development of L.
laevigata: taraxerol, lupeol, trans-melilotoside, cis-melilotoside, adenosine, patuletin 3-0-[beta]-D-glucopyranosicle, kaempferol 3-0-[beta]-D-glucopyranoside, quercetin 3-0-[beta]-D-glucopyranoside, methyl-3,5-di-O-caffeoyl
quinate and 3,3',5-trihydroxy-4',6,7-trimethoxyflavone.
Synthesis of CGA from cinnamate requires four enzymes to complete two hydroxylations on the aromatic ring and a conjugation of the hydroxycinnamate and
quinate moieties: cinnamate-4-hydroxylase (C4H), 4-coumaroyl-CoA ligase (4CL), coumaroyl-CoA:quinate hydroxycinnamoyltransferase (CQT), and coumarate/coumaroylquinate-3-hydroxylase (C3H).
Among the 18 MFS genes, 11 genes belong to the SP family (Table S11), which is the largest MFS family transporting sugars such as glucose, fructose, mannose, galactose, xylose, maltose, lactose, [alpha]-glucoside, and
quinate [56].