Leaves glabrous to subglabrous above, tomentose beneath; basal leaves rosulate to sub-rosulate; petiolate
to pseudopetiolate; blades elliptic, ovate, obovate, to deltoid, margin entire to lyrate, pinnately to palmately veined; upper leaves similar to the basal ones but few and reduced.
Leaves rosulate; sessile, petiolate to pseudopetiolate; blades linear-oblanceolate to orbicular, pinnately veined, margin entire to deeply pinnatisect, densely pubescent on both surfaces or only beneath.
Leaves alternate; petiolate, the petioles clasping; blades narrowly lanceolate to elliptic, coriaceous, pinnately veined, margin entire, abaxially buff-tomentose.
Leaves rosulate to spreading; sessile, petiolate
, pseudopetiolate to clasping; blades lanceolate, broadly orbicular to cordate, pinnately veined, margin usually spinulose, entire to lyrate, glabrous to pubescent.
Leaves glabrous to pubescent; basal leaves rosulate; petiolate, sessile to clasping; blades linear, lanceolate, ovate to orbicular, palmately to pinnately veined, margin entire, lyrate to pinnatisect; upper leaves alternate, reduced; clasping.
Leaves alternate; petiolate to subsessile; blades linear, oblong to ovate, pinnately veined, margin entire to spinosedentate, glabrous to pubescent.
Leaves alternate, glabrous, sessile or shortly petiolate, the veins palmate or reduced to one, with abaxial fiber cap contacting neither epidermis.
Leaves petiolate to narrowed at base and sessile (0); leaves broadly sheating at base (1) 15.
Leaves simple and entire, normally petiolate
, rarely sessile, usually opposite, rarely alternate or whorled; stipules absent or small and caducous, almost always with toothlike colleters in axil of leaf, sometimes on petiole, in a cluster adaxially at juncture of petiole and base of lamina or along midrib above, rarely with domatia abaxially in axils of veins.