These included small seed size, a protuberant-and-tapered base with a prominent abscission zone, and a persistent perianth (seed covering).
Calyx: Perianth five-parted: leaves oblong, acuminate-obtuse, concave.
Staminate inflorescences capitate, 10-12 flowered in a single raceme, peduncle to 10 mm long, pedicels up to 1 mm long; perianth
campanulate, 6 mm long, 6-lobed, each lobe 1 mm long; stamens 6 alternate with perianth
lobes, anthers bilocular, dorsifixed, elliptic < 1 mm diameter, extrorse, generally 1 or 2 reduced to staminodes; the fine filaments connate basally, borne at the base of a pistillode 3 mm long; lower part of the perianth
reduced to 2 or 3 small involucral bracts; indumentum densely short velutinous.