What is still poorly understood is the position of the many illustrated laciniate
pinnule forms within the frond structure of M.
Cypselae truncate at the apex, glabrous to pubescent; pappus with three, rarely two or four, narrowly paleaceous bristles, longitudinally plicate, apically plumose to laciniate
, rarely crown-like.
Ligule a membranous rim, the tip laciniate
, 1.5-2 mm long, truncate, glabrous.
Florets bisexual, zygomorphic, transitional between actinomorphic and ligulate, often villous; anther apical appendages apiculate to caudate, tails laciniate
, long, acute, tails connate; style bilobed, branches appressed, dorsally pubescent reaching the bifurcation point, hairs acute.
Sessile, cauline leaves (sometimes improperly called "ligules" on Gunnera): (0) absent; (1) like other leaves, but smaller and sessile; (2) small, straplike; (3) small, shieldlike; (4) budlike; (5) pinnatifid to laciniate
," but no plant matching both the type (leaf)