Genitalia: Dorsal labiate
plate without sclerotized rings, with two distinct sclerotized bands, spermathecal gland placed near posterior margin (Fig.
Epigynum with a wide labiate
median lobe and a pair of epigynal teeth.
Female: Sclerotized rings of dorsal labiate
plate moderately sized, folded (Fig.
Reynolds (eds.), Advances in labiate
science, Royal Botanic Gardens, Kew.
5a, b): Sclerotized rings strongly folded basally, outer margins thickened, distally compressed; dorsal labiate
plate spiculate on outer margins of sclerotized rings; ventral 1ablate plate spiculate more broadly across sclerotized rings; opening of common oviduct moderately sclerotized; inter-ramal sclerites suboval, rounded apically, spiculate basally; inter-ramal lobes claw-shaped, narrowed distally, apex rounded, weakly spiculate, mostly on apex; medial portion of the inter-ramal sclerite short (Fig.
Bursa copulatrix asymmetrical, with heavily sclerotized projections extending from anterior margin of dorsal labiate
18): Bursa copulatrix with fields of spinules in and around sclerotized rings; dorsal labiate
plate continuous and distinct; sclerotized rings large, oval, rather thick-rimmed.
2I.): sclerotized areas present on lateral margins of rami between dorsal and ventral labiate
plates; rings weakly sclerotized.
The dorsal labiate
plate is medially depressed to receive the common oviduct.
10): Dorsal labiate
plate of bursa copulatrix broad and rather short, sclerotized rings small or medium-sized, set far apart from each other, vestibulum membranous, posterior margin of fold within bursa copulatrix weakly sclerotized, valvifers 1 weakly sclerotized, posterior wall membranous with roughly symmetrical, paired, elongate oval sclerites.
11): Posterior wall a simple sclerotized plate: sclerotized rings moderately infolded laterally with the lateral portion of the underlying dorsal labiate
plate triangular plate: vestibulum relatively short and simple.
4); and the accessory sclerite on the lateral portion of the dorsal labiate
plate laterad to sclerotized ring (Fig.