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The possible exceptions, histidinal and histinidol, may have resulted from components of a vestigial cofactor pathway.
Referring to Figure 1, all compounds are derivatives of carboxylic or phosphoric acid or both, with the exception of histidinol and histidinal. These exceptions, as noted above, may be part of a cofactor pathway that was incorporated into an amino acid in order to place the imidazole functional group in proteins (unpubl.