Researchers have illustrated that geraniol can be synthesised by geraniol synthase from geranyl pyrophosphate
(GPP) [10,15], then by a dehydrogenase [10,19] or an oxidase [10,17] be converted to geranial.
The transient intermediate (10) undergoes an El elimination to generate a new carbon-carbon double bond and form geranyl pyrophosphate
Through painstaking biochemical analysis, the researchers discovered that this enzyme converts the substrate geranyl pyrophosphate
into linalool, a volatile compound with what Pichersky describes as a "wine-sweet" smell.