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The virulence of the pathogenic biotype 4/O:3 is attributed to the presence of chromosomal and extrachromosomal genes. The plasmid for Yersinia virulence (pYV) encodes the adhesin A (YadA), the Yersinia outer proteins (Yops) from type III secretion system, and the transcriptional regulator gene (virF) 7, 8].
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