Telia amphiphyllous, small, rounded, scattered or in circular groups, dark brown to blackish, erumpent or covered by epidermis.
UREDINIA abaxial, distributed over the whole surface, small, occasionally in large irregular groups, pulverulent, slightly erumpent, reddish brown in color.
Telia hypophyllous, in groups on orange pustules 3-8 mm diameter, sometimes following veins, infection sometimes causing distortion of apical leaves, erumpent
, hemispheric, yellow-brown, 200-300 pm diameter, surrounded by host tissue.
From the mass of host cells in which parasite nuclei and mitochondria replicate, an erumpent
thallus differentiates, protrudes from the host, and ultimately produces reproductive structures (spores, gametes).
Infection sites were evident as small chlorotic flecks on the leaf, and when these developed a distinct orange color and the overlying epidermis became erumpent
, the uredinium was considered formed.
Hysterothecia gregarious, erumpent
, superficial, straight to flexuous, irregular in shape especially when crowded, 0.