Once the scutella are yellow or brown, they may dehisce
. When a raindrop hits the cup-shaped surface of the scutellum it inverts and falls off the scale and the mechanical force of this event causes the scale to throw the mature fruits.
All mature fruits were collected as they began to dehisce
. This occurred on 17 July at Vine, and on 12 August at Cincinnati.
Some of the capsular fruits open completely to shed the seeds, sometimes with individual seeds adhering to the valves as they shed; others remain attached to the twig and dehisce their seeds as the fruit is shaken by the wind.
f., with large trees native to western Malaysia, Sumatra, Borneo, is an exception, having 3-winged fruits that dehisce into three valves as well illustrated by Leenhouts (1956).
Nuts are enclosed in a fibrous, four-valved husk that dehisces at maturity in September to December.
The pericarp dehisces along one side of the nut when mature in midsummer.
These cupules are usually armed with spines but dehisce at maturity to expose and release the nut.