Leaves opposite, rarely whorled, often with cluster of colleters adaxially
at base of blade (distributed along the length of midrib in some species of Mandevilla), sometimes with domatia abaxially in axils of secondary veins with the midvein.
C"-shaped trace: 0 = absent; 1 = abaxially convex; 2 = adaxially
1 cm wide near the base, green to reddish toward apex, opaque, inconspicuously white-lepidote abaxially with trichomes disposed along the intercostal zones and not obscuring the color of the blades, adaxially
glabrescent toward the apex, distinctly nerved on both sides, apex long acuminate-caudate, pungent, margins glabrous, sparsely spinose, spines 0.
This taxon also differs from the other members of Section Ormopteris by the slightly sulcate axis adaxially
Plants terrestrial; rhizomes erect to decumbent; petioles scaly toward the base, with 4-10 vascular bundles, the two adaxial bundles enlarged; laminae 1-pinnate-pinnatifid to 4-pinnate-pinnatifid, catadromic above the basal pinnae; basal pinnae inequilateral and more developed on the basiscopic side or (less commonly) equilateral; rachises, costae, and costules not grooved or only shallowly so adaxially
, scaly and pubescent abaxially, densely pubescent on the adaxial surfaces, the hairs whitish, spreading to antrorsely strigose, multicellular, if glands present, these ca.
Indument on costae and veins adaxially
, margin of segments, rarely on laminar tissue adaxially
of spreading acicular trichomes mostly 0.
5 cm, thinly coriaceous, strongly nerved mainly adaxially
, inconspicously and sparsely white-lepidote to glabrescent, green, distinctly canaliculate mainly toward the base, apex acuminate-caudate, recurved.
M1 separated from M2 by a distinct furrow slightly inclined anteriorly, then strongly curved forward adaxially
Its species differ from others in the genus by having a distinctive type of hair within the grooves of the rachises and costae adaxially
3m, n), each subtending an ovuliferous scale consistently bearing two adaxially
placed ovules, their micropyles directed toward the cone axis.
grooved, 8-18 mm long, glabrous to lightly puberulent, petiolar glands absent.
5-10 cm, inconspicuously lepidote, green toward the apex, strongly coriaceous; blades narrowly subtriangular-lanceolate, not narrowed at the base, 15-24 x 5-6 cm, green, inconspicously and sparsely white-lepidote mainly adaxially
to glabrous, nerved, suberect with recurved apex, apex acuminate.