Frontoparietals thin; anterior margins extending laterally over dorsolateral optic margin of sphenethmoid, but well behind posterior margin of nasals; in contact medially; posterolaterally in contact with epiotic eminences; not elevated posteriorly to form crest.
Prootics and exoccipitals fused; entire posterior end of skull well ossified; optic foramina ventrally marginated by cartilage; oculomotor foramina marginated anteroventraly by cartilage; prootic and jugular foramina with bony margins; with small bony knob formed below epiotic eminences above occipital condyles.