Sharpey's fibres, which were for the attachment of the scale in the skin, were present already in early stages of the scale development.
The density of Sharpey's fibre tubules in the middle and basal layers can be high (Figs 813; 913, F); the fibre tubules are distributed irregularly or relatively horizontally.
Sharpey's fibre tubules occur in both the middle and the lower layer.
Vascular canals and Sharpey's fibre tubules occurred in both the middle and lower layers.